Hypothetical distribution patterns of the meadow brown Maniola jurtina in Europe during the last glacial maximum (dark hatched areas). Postglacial expansions are indicated by solid arrows. The postulated actual hybrid zone is shown by the hatched area. Question marks indicate lack of information concerning ancient distribution patterns in the South and present distribution of hybrid populations in the North. Redrawn from Schmitt et. al. ; based on Thomson  and Schmitt et al. .
Schmitt Frontiers in Zoology 2007 4:11 doi:10.1186/1742-9994-4-11